Description

Key Characters:

Growth Form: Perennial herbs.

Stems: Producing long stolons.

Roots: Stout scaly rhizomes usually 1–4 cm long.

Leaves: Leaves compound (odd-pinnate). In a basal rosette. Blades obovate–cuneate, leaflets 2.1–5(–6) cm long, 1.2–3.5 cm wide, lateral petiolules 0.2–0.8 cm long, the terminal ones 0.2–1(–2.5) cm long. Upper surfaces glabrous or with a few scattered long hairs; lower surfaces densely silky long–strigose, blades, somewhat coriaceous. Margins coarsely serrate, sometimes only in upper ⅔. Upper surface with conspicuously impressed veins. Petioles (3–)12–16(–21) cm long, densely long–strigose. Stipules brownish, membranous, 1–1.6 cm long, coherent to base of petioles.

Flowers: Flowers 1 to several in scapes usually ca. ½ as long as petioles at anthesis, apparently elongating after anthesis to nearly as long as petioles. Flowers bisexual (perfect) or unisexual (and then plants polygamodioecious). Pedicels usually 20–50 mm long; epicalyx lobes narrowly lanceolate, lobes alternate with the calyx lobes and persistent, (3–)5–6 mm long, densely silky long–strigose. Calyx of 5 sepals, lanceolate, distinct, imbricate, ca. 7–8 mm long, enlarging to 9–10 mm long in fruit, densely silky long–strigose. Corolla of 5 petals, white, broadly obovate, distinct, imbricate, 8–11 mm long. Stamens numerous, in 1–3 whorls, persistent; filaments slender; anthers dithecal. Ovary superior, carpels numerous, attached to the elevated conical receptacle; styles lateral.

Fruit: Fruit an aggregate consisting of numerous minute achenes borne on the surface of the enlarged fleshy receptacle; red; broadly conical; ca. 1–1.5 cm long; hirsutulous. Achenes ca. 1.5 mm long; slightly embedded in shallow pits of the receptacle. Seeds 1 per achene.

Ploidy: 2n = 56*

Habitat: Dry; disturbed areas; wet forest; and subalpine shrubland; and coastal.

Elevation Range: 1,160–3,070 m.